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The Ark is the location where the events of the Act V - Ascension take place. It is Galath's secret hideout and it contains high-tech gear (such as elevators and holograms). Occasionally, a dragon may appear at the path leading to the Ark. Ryth can also be encountered at the first floor, after you open the golden treasure chest, and have a Solar Transport weapon.
Note: Worker's rocket which is seen in the final battle is also named the Ark, as mentioned by the Guardian of Heaven in the observatory (requires Map of Heaven).